Semple, J. C. & J. G. Chmielewski. 1987. Chromosome number determinations in fam. Compositae, tribe Astereae. II. Additional counts. Rhodora 89: 319–325.
Semple, J. C. 1984. Cytogeographic studies on North American Asters. I. Range surveys of Virgulus adnatus, V. concolor, V. georgianus, V. grandiflorus, V. novae-angliae, V. oblongifolius, V. patens and V. walteri. Amer. J. Bot. 71: 522–531.